Biology Courses

Note that this page has two tabs on it. Clicking on the left-hand tab will yield links to web sites for current-semester biology courses. Clicking on the right-hand tab will yield descriptions of all Biology courses.

You can filter the list of course descriptions below to show different subsets of Biology courses. By default (with all of the drop-down boxes set to "Any"), you will see all courses. Choosing other options in the drop-downs limits the list of courses. For example, if you choose "Evolution and Biodiversity" in the Core Area box, and "Yes" in the Lab box, you will see descriptions of all courses in the Evolution and Biodiversity category that meet the lab requirement.

First semester of a full year course for science majors. Lecture: introduction to biochemical basis of living systems, cell and molecular biology, mitosis and meiosis, principles of genetics, developmental biology. Lab: selected exercises on cell structure, enzymes, photosynthesis, mitosis, genetics and embryology. Lecture: hour exams and final. Lab: quizzes, report, practical.

Second semester of a full semester course for science majors. Lecture: Topics in organismal and evolutionary biology: evolution; the diversity of life; plant and animal structure and physiology, ecology. Lab: plant anatomy and development; animal diversity; excretion, rat dissection, ecology. Hour exams, final; lab quizzes and reports; lab practical. Prerequisite: Biology 100 with grade of C or higher.

Lecture: The cell biology, genetics, physiology, evolution, and ecology of animals. Lab: survey of animal groups; experiments in animal structure and function including dissection. Two multiple-choice hour exams, final; many lab quizzes and writing assignments; lab practical exams. Not for Biology major credit.

An introduction to plant biology with emphasis on the flowering plants. Selected topics from anatomy, cell biology, evolution, genetics, molecular biology, morphology, physiology, and systematics. Labs illustrate principles discussed in lecture and include a survey of the plant kingdom. Lecture exams, lab quizzes, and lab practical. Not for Biology major credit.

Introduction to biological principles underlying some current social problems. Designed to provide non-biology majors with the basic scientific knowledge that an informed citizen requires to develop thoughtful positions on sometimes controversial questions related to medical ethics, environmental degradation, biotechnology and education. Not for Biology major credit.

Physiology of the major systems of the human body in health and disease. Intricacy of the human body; good health practices. Collateral issues include birth control, sexually transmitted diseases, nutrition, and health risks associated with drugs and smoking. For nonscience majors; not for Biology major credit.

This course explores the evolution, extinction, and conservation of biodiversity on earth. We will survey the diversity of both ecosystems and organisms found throughout the globe. The generation of biodiversity will be explained by both ecological and genetic approaches to evolution. The loss of biodiversity due to historic extinctions and current human activity will be examined. We will use examples from all over the world, but will focus on many examples from New England and Massachusetts. One section will survey the history of wildlife in Massachusetts since European colonization. Not for Biology major credit.

Team-taught course with faculty from Departments of Physics, Astronomy, Geosciences, Microbiology, Biology, Anthropology, Psychology, Computer Science, Philosophy, and History. Lecture topics include Cosmic Fundamentals: space, time and relativity; matter, energy and the Standard Model. The Universe: cosmology; dark matter and dark energy; origin and evolution of galaxies, stars, the elements, the solar system, and the Earth. Life: its nature and origin; the microbial world; origin and evolution of plants and animals. Intelligence: behavior and the mind; origin and evolution of biological intelligence; artificial intelligence and robotics; life and intelligence in the universe. “Friday Forum” topics include Albert Einstein: “Indisputably One of the Greatest Men of Our Time” –George Bernard Shaw; Quantum Weirdness: “Nobody Understands Quantum Physics” –Nobel Laureate Richard Feynman; Quantum Gravity and String Theory and the Anthropic Principle and the Multiverse; Meteorites: Messengers from Space; Paradigm Shift: From a Fixed Crust to Continental Drift and Plate Tectonics; Charles Darwin: “Most Important Person Who Ever Lived” –Nobel Laureate James D. Watson; Extremophiles: Life on the Edge; Debate: Are Viruses Alive?; Debate: What Is the Mind?; Debate: Will Artificial Intelligence and Intelligent Robots Ever Surpass Human Beings?; Debate: Is Extraterrestrial Intelligence Common? and The Grandest Finale: How Will It All End? Two lectures/week and a “Friday Forum” with faculty-faculty, faculty-student, and student-student discussions and debates. Not for Biology major credit.

An introduction to the workings of the cell, focusing on themes of cellular structure, dynamics and energetics. This course is intended for students interested in a broad interdisciplinary approach to the biological sciences: frequent connections to chemistry, physics and mathematics will be made as the cell, its inner workings and malfunctions, are explored. In the laboratory, students will work in teams to conduct multi-week inquiry-based experiments in a laboratory 'core facility' to complement and expand on the lectures. This first semester course is prerequisite for the second semester in a full year introductory course sequence for life science majors.

Process of biological evolution and the evolutionary history of life on earth. Historical framework of evolutionary thought from pre-Darwinian times to Darwin and through the development of the modern evolutionary synthesis (Neo-Darwinism). Major features of biological evolution including microevolution (the evolution of biological populations), speciation (the origin of species), and macroevolution (evolution above the species level). Biological systematics, including phylogeny reconstruction and classification. Cosmic or inorganic evolution-- the origin and evolution of the physical universe, including evolution of the elements, stars, the earth's crust, atmosphere, and oceans. Chemical evolution and the origin of life. Geologic time and the fossil record. Microbial, plant, and animal evolution. Primates and the evolution of humans; cultural evolution. Origin of major evolutionary innovations stressed, including evolution of photosynthestic, oxygen-releasing bacteria, nucleated cells, sexual reproduction, true roots and leaves, woody plants, the seed and flowers, animals with a true body cavity, insects and their societies, and the vertebrate jaw, the transition from fins to limbs, the evolution of the shelled land egg, vertebrate flight and warm bloodedness, and bipedalism and the human mind. Prerequisites: a grade of C
or better in Biology 100 and 101.

Introduction to genetics including classical, cytological, molecular, biochemical and populational aspects, in a wide variety of organisms. Quizzes, hour exams, final. Prerequisite: A grade of C or better in Biology 100 and 101; completion of Biology 285 recommended.

The basic objective of this laboratory is to introduce the methods and pleasures of genetic investigations, using a variety of organisms. It also reinforces Introductory Genetics (Biology 283). Topics include: Mendelian genetics, gene maps, variation in both DNA and in proteins, mutation induction and selection and DNA polymerase chain reaction. Prerequisite: BIOL 283 (may be concurrent).

Course designed for sophomore-level majors in life sciences. Building upon concepts introduced in BIOL 100/101, consideration is given to structure and function at the cellular, subcellular, and molecular levels. The course is equally divided between aspects of molecular and cellular biology. Prerequisites: a grade of C or better in Biology 100 & 101.

A course in general ecology designed for undergraduate majors in biology. The course will cover the following topics: how the world works, its structure, history, and evolution; the Earth in space and extra-terrestrial influences; the energy budget and atmospheric circulation (weather); ecosystems and the flow of energy; biomes of the Earth; biogeochemical cycling; adaptations of plants and animals to their environments; population dynamics; interactions between organisms including the concepts of symbiosis and succession; human technology and ecological problems; and ideas for developing new relationships between human technology and ecological problems; and ideas for developing new relationships between humans and the natural systems we need for future survival. Prerequisite: a grade of C or better in BIOL 100 &101 or 102 & 103.

Lectures cover the physiology of humans and other vertebrates on a system by system basis (e.g. circulatory system, respiratory system, digestive system, etc.). Emphasis is placed on understanding fundamental physiological concepts such as diffusion, membrane potentials, biomechanics and biocontrol. Problem sets and exams give students practice working with physiological concepts. This course concentrates primarily on human physiology, but examples from other vertebrate animals are used to illustrate some physiological phenomena. Prerequisites: Grades of C or better in BIOL 100 & 101.

A lecture series introducing the natural history, reproductive biology, functional morphology, and evolution of the major groups of marine vertebrates. Species native to the North Atlantic and Caribbean will be emphasized, including selected sharks, teleosts, turtles, birds and mammals. Grading will be based on two multiple choice exams. A note set must be purchased (at cost), but no other text is required. Prerequisites: grades of C or higher in Biology 100 & 101.

Please click through to view descriptions of different instructors' versions of the writing course.

In this class we will discuss concepts and applications of modern DNA technology including an introduction to the basic concepts pertaining to the emerging field of genomics. We will begin by describing key molecular methods (cloning, sequencing, blotting, PCR) and how they are used in gene analysis. We will then move on to consider how entire genomes are analyzed, and will familiarize ourselves with some of the basic bioinformatics' tools that are commonly used by working biologists. Finally we will consider the methods used to manipulate genomes as a means to determining gene function. This course is intended for sophomores and juniors, and should serve as a bridge between 200-level courses and more advanced, specialty courses (e.g., 500-level courses). Prerequisite: Biology 285 or Biochem 285.

We have two goals in this course. The first, and most important, is to introduce undergraduate Biology students to some of the many fascinating aspects of Plant Biology, especially as these differ from animal biology. For instance, did you know that plants are moving (on a large scale) all the time? It’s the truth, but in a very different time scale than we animals use. How do plants do that without the benefit of muscles and skeleton? Have you ever thought about how, in the absence of a pumping heart, plants’ circulatory systems work? After all, the water at the top of a tree got there from roots in the ground, but no pump was involved. Plants don’t have an immune system, and yet, they ‘stand and fight’—literally rooted to the spot—taking on all types of pathogens, as well insects and other predators. What strategies do plants use to overcome these attacks? Have you ever wondered about how biotechnology is used in agriculture? We have all heard news stories about GMO’s (genetically modified organisms). What are these and what makes them useful or dangerous? These are the types of topics we will be covering in this course. The second goal for this course is to provide a convenient way for UMass Biology majors to accomplish their plant biology course requirement. The course is open to any student who has successfully (with a C or better) completed the Introductory Biology series Biol 100 and Biol 101.

This fundamental ecology course emphasizes the quantitative skills needed to understand and conduct field research. The lectures introduce major ecological concepts, local vegetation types, and methods and techniques of gathering and analysing data. In laboratories, students collect original data at sites in the Connecticut Valley and write an original scientific paper. Prerequisite: an introductory biology course or consent of instructor.

Learn the vascular plants of the region in their natural habitats through field trips and in the laboratory with the use of botanical keys and manuals. Field experience will include some collecting and pressing of specimens. The class also visits the herbarium and greenhouses. Recognition of certain plant families and familiarity with terminology will be gained. Prerequisite: Introductory Biology or consent of instructor.

The goal of this laboratory course is to explore how researchers address modern biological questions through the use of model organisms. The course will be taught by a team offaculty whose own research employs these model systems to answer a diverse range of biological problems, including molecular evolution, plant development, embryonic development and population genetics. Students will be introduced to several different model organisms that may include representative bacterial, plant, invertebrate, and vertebrate species. Lab exercises will employ sophisticated, state-of-the-art molecular methods and will tackle a variety of current biological questions. Prerequisites: Biology 283 or 285.

In this interdisciplinary course, we will explore the topic of imaging biological material, beginning with optics and basic microscopy. Students will perform hands-on exercises in the use of the light microscope, digital cameras, and image processing and quantification. Common pitfalls in imaging biological samples will be covered. Students will perform experiments to test and quantify various aspects of cell migration, cell cycle regulation, mitosis and endocytosis. Using the methods learned in the first portion of the class, students will design and complete a hypothesis-based experiment of their own design and present their findings. Bioimaging is a laboratory-based course. Enrollment is limited and requires permission of the instructor.

This course will cover the cell biological aspects of several plant cellular processes, including cytokinesis, cell expansion, tip growth, cell-to-cell communication, and intracellular protein sorting. An emphasis will be made on experimental approaches used to understand these processes at the molecular level. A discussion of model organisms and cell types will be included. Formats will include lectures, discussions, and in-class student presentations. Prerequisite: Biology 285.

Vertebrate collections are indispensable to the teaching of all whole organism biology courses. They are the repositories of Study of DNA, study skins, hides, preserved specimens, and comparative osteological material. Collections contain regional and global reference collections, provide secure storage for specimens of regional and historic importance, support a library of DNA samples and the specimens from which they were collected and also serve as the repository for type specimens. Maintenance of biological collections not only requires space and money but also require a highly dedicated and trained staff in order to be useful. Without constant maintenance even the smallest collections will deteriorate to the point of uselessness. Today many collections are being orphaned and/or discarded by colleges and universities. Qualified preparators, collections managers and curators are few and far between. During the course of this semester we will learn how to legally obtain, prepare, catalogue, and store new specimens and maintain an historic collection of vertebrate specimens. There will be one weekly lecture and at least 3 additional hours of lab per week. Lectures meet for 2.5 hrs. (6:00 ¬ 8:30 PM).

Honors course in Tropical Biology. Introduction to the ecology, behavior, taxonomy, and physiology of tropical organisms, with emphasis on close observation of living organisms in nature. Includes hands-on investigation of coastal and marine ecosystems. Students will participate in a field trip to Costa Rica during the Spring Break. Prerequisite: permission of instructors.

Structure and function of components of the plant cell, including the wall, membranes, vacuoles, the cytoskeleton and various organelles. Aspects of development at the molecular, tissue and whole plant level. Current theories pertaining to how plants react to hormones, light and daylength. Responses to stresses such as drought, temperature and touch, and the nature of plant defenses against predation and disease. Prerequisite: Biology 103 or Biology 100 & 101.

This course focuses on the processes affecting the distribution of genetic variation in populations of organisms, through space and time. The processes studied are the ones that operate during evolutionary change. Topics covered will include the Hardy-Weinberg principle, gene flow, genetic drift, recombination and linkage disequilibrium, natural selection, the effect of mating systems on diversity, and the neutral theory of evolution. Examples illustrating key concepts will be drawn from various kingdoms of life. The course will consist of lectures and occasional in class discussion. Prerequisites: Biology 280 or equivalent; Math 127 or Statistics 240 or equivalent.

Detailed approach to the structure and evolutionary relationships of vertebrates. Lecture: evolutionary and functional significance of structures in different groups. Lab: evolutionary trends and specializations, experience in dissection. 2 hour exams, final; 2 lab exams. Prerequisite: Biology 100 & 101.

Introduction to vertebrate history emphasizing fossil forms. Topics include: changes in locomotory, feeding, and defense structures, modes of life of extinct animals, such as dinosaurs, faunal changes over time, and relationships among the various groups of vertebrates. Lectures at Amherst College Pratt Museum, with study in lab of display and other fossil specimens. Requirements: 2 hour exams, final, lab quizzes. Prerequisite: introductory course in biology, geology, or physical anthropology.

The relation of cell, tissue, and organ microscopic and submicroscopic structure (mammals primarily) with function. Cell fine structure and function; microscopy; tissues (epithelial, connective tissue, nerve and muscle) and selected organs (lymphatic, endocrine and other glands, circulatory, and digestive). Lab includes light microscope identification, related electron micrographs, introduction to microtechnique, demonstrations in the Electron Microscopy and Image Analysis facility. 3 exams, multiple quizzes, final; 2 1-hour lab practicals. Prerequisite: Biology 100 or equivalent.

An advanced course for students who have already taken an introductory course in evolution and who are willing to make an active contribution to classroom discourse. We will discuss both evolutionary mechanisms and evolutionary history. Potential topics include evolutionary genetics, the role of chance in evolution, speciation and species concepts, the origin of life, the tempo of evolution, extinction, the evolution of behavior, evolutionary history of selected groups, research methods in evolution. Prerequisite: Biology 280 or equivalent course.

The course provides an overview of the systematics, anatomy, and evolution of all the major, living lineages of amphibians and reptiles, with an emphasis on the herpetofauna of Eastern North America and New England. The laboratory is organized around three approaches: anatomical studies; studies of live organisms; and studies of regional and global amphibian and reptile diversity. If weather permits, there will be one field trip near the end of the semester. Some dissection is required. Prerequisite: Biology 521 or permission of the instructor.

This is an introductory course designed to familiarize students with the diversity of fishes. We will provide an overview of the biology, evolution and ecology of fishes. A phylogenetic approach will be used to look at major primitive to advanced fish groups. No prior coursework is required to take this course, but students are expected to have a general biology background and be enthusiastic in learning about this diverse group of organisms. The textbook to be used is “The Diversity of Fishes” by Helfman, Collete and Facey 2004. Only selected portions of the text will be required during the course. The lab is designed to supplement the lecture course with hands-on dissection, anatomy of preserved specimens and dry skeletons and identification of major lineages. 1 essay exam, final, 2 lecture quizzes, 2 lab practicals.

Lecture: origin of birds, speciation, diversity, flight, territoriality, migration, navigation, communication, conservation. Lab: bird identification, anatomy, field studies. Text and field guide required. Lab practicals, 2 lecture exams plus final. Prerequisite: upper level biology course or consent of instructor.

Lectures and readings on comparative biology, phylogenetic relationships and evolution of mammalian groups. Lab involves a detailed introduction to the New England mammals and study of selected representatives of most mammalian orders, emphasizing systematics, function and morphology. 2 hour exams, 2 lab exams, and final. Prerequisite: any life science course beyond the introductory level.

Animals have evolved a remarkable diversity of behavioral patterns, used in a wide range of ecological and social contexts. Our first goal in this course will be to examine the mechanisms responsible for the expression of behavior: for example, how do birds locate prey; how do crayfish avoid becoming prey; and how to crickets and birds develop species-specific communication signals? To help answer these questions we will make use of neurobiological, hormonal, genetic, and developmental perspectives. Our next goal in the course will be to examine the evolutionary bases of behavior, asking for example why animals move, forage, hide, communicate, and socialize as they do. To address these questions we make use of optimality theory and other behavioral ecological perspectives. Other topics in the course will include sexual selection, human behavior, and the role of behavior in establishing biodiversity. Prerequisite: introductory biology or psychology course; or consent of instructor and at least sophomore level standing.

Discussion of cell structure and function; emphasis will be placed on the properties of individual molecules that contribute to cell function. Topics will include the mechanism and regulation of cell division; interactions of cells with each other and with the extracellular environment; cell motility; and the organization of membrane systems. Techniques used to study cells will also be discussed. Format will include both lectures and class presentations; quizzes, mid-term exams and written assignments will be included. Prerequisite: Biology 285.

Physiological principles governing the function of major organ systems (nervous, circulatory, respiratory, endocrine) and their interactions in vertebrates emphasizing mammals especially humans. Lab exercises designed to illustrate physiological principles using modern approaches. Prerequisite: Biology 285 or equivalent and at least one semester of Organic Chemistry.

Lectures cover the physiology of vertebrates and invertebrates on a system by system basis (e.g. circulatory system, digestive system, etc.). Comparisons between animals within each system and adaptations to "extreme" environments are emphasized. Weekly problem sets provide practice in physiological reasoning for each system covered. Animal design projects involve modeling the physiological systems of an extinct animal. Prerequisite: a grade of C or better in Biology 100 & 101.

The role of hormones in growth, metabolism and reproduction, molecular mechanisms of hormone action, and feedback control of hormone secretion. 2 hour-exams, final, and 1 class presentation. Prerequisite: Biology 297A or consent of instructor.

Biology of nerve cells and cellular interactions in nervous systems. Lectures integrate structural, functional, developmental, and molecular approaches. Topics include neuronal anatomy and physiology, membrane potentials, synapses, development of neuronal connections, visual system, control of movement, and neural plasticity. Text: Essentials of Neural Science, Kandel et al.; reserve readings, 2 hour-exams, final, short critique paper. Prerequisite: Biology 285 or 523; or both Psychology 330 and Biology 100.

Analysis of organismal development, with special attention to cell-cell interactions, cells fate determination, gene regulation, signal transduction, pattern formation and terminal differentiation. The emphasis will be on molecular approaches to these problems. Prerequisites: Biology 285 or equivalent recommended.

This course will explore animal communication from several biological perspectives. We will explore how animals use different modalities of communication (sound, smell, electricity, etc.) and how these modes of sending and receiving information are limited by environmental constraints and their functions. We will look at the physiological and anatomical aspects of signal production and perception. The class will discuss the different types of messages encoded in signals and how they evolved. We will explore the evolution of sexually selected forms of communication (antlers, bird song, etc.) and the theories that attempt to explain their function and evolution. The lectures/discussions will draw on examples from a diverse selection of animals (insects, fish, birds, and mammals). Students will also work on projects where they will learn how to analyze and interpret different forms of vocal and visual communication.

Methods in data capture and analysis with an emphasis on using and developing approaches to specific biological problems. The R computing environment will be used as a way of accessing libraries of traditional approaches as well as developing novel approaches to biological data analysis including time series, morphometrics, DNA and protein sequence data and experimental design. Prerequisite: Math 127, 128 or equivalent.

Earth's environment from Archean times to the present; the planetary impact of the origin and evolution of life. Microbial communities, metabolic and cell evolution. International faculty accessible via the interactive lecture-electrowriter system. Class discussion and oral presentation. Limited to senior science majors.

Simultaneous enrollment in Biology 597M (lab) is required. An advanced course focused on the evolution of macromolecules and the reconstruction of evolutionary history of genes, proteins and organisms. Potential topics include databases and sequence matching, molecular phylogenetics, gene duplication and divergence, genome evolution, and horizontal gene transfer. The course will consist of lectures, computer demonstrations and class discussions. Text: Molecular Evolution, W.-H. Li, 1997 and readings from primary literature. 2 hour-exams and 1 term paper. Prerequisite: Biology 280 or equivalent course.

This course covers current topics and advanced concepts and techniques in genetics and their use in answering fundamental questions in biology. Theoretical background and experimental approaches will be emphasized. Topics will include, but are not limited to, gene and genome structure and function, tools and approaches of genetic analysis, recombination and mapping, and developmental and quantitative genetics. Prerequisite: Biology 283 with grade of C or higher.